Penguins may look charmingly awkward on land, but new research shows their bodies are finely tuned for powerful, efficient movement both on land and underwater. A team of anatomists from Midwestern University, in collaboration with SeaWorld San Diego and Scarlet Imaging, has published a detailed study of the limb musculature of the macaroni penguin (Eudyptes chrysolophus), uncovering previously unknown features that help explain how these birds swim, dive, and even stand upright.